He NP, Liu CC, Piao SL, Sack L, Xu L, Luo YQ, He JS, Han XG, Zhou GS, Zhou XH, Lin Y, Yu Q, Liu SR, Sun W, Niu SL, Li SG, Zhang JH, Yu GR*. Ecosystem traits linking functional traits to macroecology. Trends in Ecology and Evolution, doi: 10.1016/j.tree.2018.11.004.论文链接：https://www.sciencedirect.com/science/article/pii/S0169534718302751

“Here, ecosystem traits are traits or quantitative characteristics of organisms (plants, animals, and microbes) at the community level expressed as the intensity (or density) normalized per unit land area ”，